Gyeltshen, K. (2018). "Population status and diet of sympatric hornbills in Jomotsangkha wildlife sanctuary(JWS), Bhutan''. Forest Information Division and Forest Research Institute.
शिकागो शैली (17वां संस्करण) प्रशस्ति पत्रGyeltshen, Karma. "Population Status and Diet of Sympatric Hornbills in Jomotsangkha Wildlife Sanctuary(JWS), Bhutan''. Dehradun: Forest Information Division and Forest Research Institute, 2018.
एमएलए (9वां संस्करण) प्रशस्ति पत्रGyeltshen, Karma. "Population Status and Diet of Sympatric Hornbills in Jomotsangkha Wildlife Sanctuary(JWS), Bhutan''. Forest Information Division and Forest Research Institute, 2018.
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